According to Official Guinness Records,
Created in 2001 by H.J. Tsai, a professor of fisheries science at National Taiwan University, subsequently sold by the Taipei-based Taikong Corporation, and dubbed Frankenfish, the world’s first man-made bioluminescent fish are green-glowing specimens of the zebra fish, a popular aquarium species, whose bioluminescence is due to the introduction of jellyfish DNA. Officially known as night pearls or TK-1, they were followed in 2003 by a second artificial strain of bioluminescent zebra fish, the TK-2, this time glowing red rather than green, having received a gene for red bioluminescence from a species of red-glowing coral.
For a complete list of 2001 records, please visit 2001 Guinness Records in Taiwan.